Fossil evidence for alleged apemen—Part 2: non-Homo hominids
by Peter Line
The aim of this overview is to critically analyze the fossil evidence for the alleged
apemen that are not classified in the genus Homo. Each of these hominid
candidate species will be examined in roughly ‘evolutionary’ chronological
order, beginning with the earliest. The skeletal morphology of these alleged apemen
indicates they were extinct apes that had nothing to do with human evolution.
M.P.F.T. <www.cnrs.fr/cw/fr/pres/compress/Toumai/
Tounaigb/illustragb.html>
Sahelanthropus tchadensis, best known by the cranium (TM 266-01-060-1)
nicknamed Toumai, was announced to the world as the earliest-known hominid in July,
2002, amid extraordinary publicity. However, the relegation of Sahelanthropus
to an ape, by some experts, did not receive the same media attention.
The supposed apemen belong to a category of fossils known as hominids, a group that
includes living humans, their ancestors and any other ape-like creature believed
to be more closely related to humans than to chimpanzees. Some evolutionists now
refer to this group as hominins.1
The term hominid is nonsensical from a creationist point of view, because the hominids
were either human or apes, not anything in-between. The term is used here only to
categorize these fossils, and does not imply a belief that such creatures really
existed. According to evolutionists, the defining moment for determining a hominid
is the hypothesized split between humans and chimpanzees. In the past the human
and chimpanzee calibration point has been determined by a molecular clock to be
5 million years ago (Ma), using as calibration point the supposed split between
Old World monkeys and apes 30 Ma, as determined by conventional radiometric dating
of fossils.2 The split is now thought,
by evolutionists, to have occurred about 5–7 Ma. Radiometric dating3 and molecular clocks4
are based on unproven and problematic assumptions. When the evolutionary ages of
fossils are given in this article, it is for the purpose of putting them in an evolutionary
context, and it in no way implies that these age dates are valid.
The australopithecines refer to the members of the genus Australopithecus,
and in earlier times this was the only hominid genus apart from Homo. However,
in recent years many hominids have been assigned to new taxa outside Australopithecus,
such as the genus Ardipithecus, Sahelanthropus, Orrorin
and Kenyanthropus. From a creation point of view, it should be remembered
that a species is not equivalent to a biblical kind.5
Sahelanthropus tchadensis
If publicity surrounding the announcement of a fossil find is anything to go by,
then Sahelanthropus tchadensis is the frontrunner for the title of earliest
hominid, as the hype was extraordinary even for apemen fossils, which have a reputation
for making a lot of ‘noise’ on arrival. The cranium of Sahelanthropus
(TM 266–01–060–1, nicknamed Toumai), found in Chad, Central
Africa, was featured on the cover of the 11 July 2002 issue of Nature,
with the title ‘The earliest known hominid’. The single cranium and
other fossils (fragmentary lower jaws) were dated by the researchers, using associated
fauna, to be between 6 and 7 Ma, with the cranium estimated to have a cranial capacity
somewhere between 320 to 380 cm3.6
The odd thing about the Toumai cranium is that it looks chimpanzee-like from the
back, but from the front it is said by one expert to pass as an advanced australopithecine.
7 According to Bernard Wood, a hominid of Toumai's supposed age ‘should not
have the face of a hominid less than one-third of its geological age’.7 Hence, if evolutionists accept Sahelanthropus
as the earliest stem hominid, then this casts legitimate doubt on the human ancestry
status of all other hominids with more ‘primitive’ faces,7
and, as such, would invalidate most of the geologically younger australopithecines.
Almost before the print had dried on the hyped headlines surrounding Toumai’s
arrival, other evolutionists, including Brigitte Senut of the Natural History museum
in Paris, expressed doubt about Toumai’s hominid status. In an interview,
she stated her view as tending ‘towards thinking this is the skull of a female
gorilla’.8 This hardly corroborates
Harvard paleontologist Daniel Lieberman’s earlier grandiose claim about Toumai
being ‘the most important fossil discovery in living memory’.9 A team led by Wolpoff argued that Toumai’s
impressive supraorbitals (large brow ridges), which are greater than in any australopithecine
or extant ape, were more likely to be a mechanical response to strain,10 and not an indicator of facial similarity
to the genus Homo. Examination of scars left on the back of the skull from
neck muscle attachments led them to conclude that Toumai was a quadruped (i.e. walked
on four legs, not two).11 After
their examination of Toumai, they stated their belief ‘that Sahelanthropus
was an ape’.12 Another
alleged hominid feature of Toumai is its small canines compared to extant apes.13 Although Sahelanthropus
differs from extant gorillas in some features, there are also many similarities,
and according to Matthew Murdock there is a strong possibility that Sahelanthropus
and extant gorillas are related in the sense that they are both members of
the same biblical kind.14
If this is correct, then the small canines may just reflect greater canine size
variation existing in this ape kind in the past, with the genetic information
for small canines subsequently lost.
Recently, claims have been made that one of the molars in the mandible was glued
in the wrong place, and that an incisor collected at the site was not featured in
the original Nature paper.15
The authors argued that the fossils attributed to Sahelanthropus ‘are
the subject of debate as to whether they represent a hominid or an ape. It is therefore
necessary to provide full and accurate details of the fossil collection.’16 As so often happens in paleoanthropology
when the establishment is challenged, the pair making the claims suffered a backlash.17 Also predictable was that
the relegation of Sahelanthropus to an ape followed the familiar pattern
of not getting anywhere near the same media attention that greeted its initial ascension
to apeman.
Orrorin tugenensis
Call
it what you like, but is it science?
|
Early in 2001 Orrorin tugenensis, claimed to have lived about 6 Ma, was
announced as a candidate for the earliest hominid (the fossil bones were found in
Kenya’s Tugen Hills), by a team of researchers led by Brigitte Senut and Martin
Pickford.18 The fossils consisted
of 13 pieces, including broken femurs, several teeth, and bits of lower jaw.19 Hence, it is not possible to know what the
head of this creature looked like. The only reason Orrorin has claims to
early hominid status, as well as claims of bipedality, is that it was found in the
right evolutionary time period. Bipedalism is considered by evolutionists to be
important evidence of hominid status. Orrorin supposedly walked on two
legs because of an alleged human-like femur (thigh bone), based on its long femoral
neck and a groove on the back of this neck which is present in humans, but absent
in chimpanzees.20 However, this
groove, where the obturator externus muscle presses against the
bone, is also found in non-bipeds,21
and according to one expert, scans through Orrorin’s femoral neck
show a chimp-like distribution of cortical bone.20
Another argument is that the head of the femur in Orrorin is proportionally
larger than Lucy’s, and evolving a large-headed femur would help dissipate
the forces caused by bipedalism.22
The team controversially suggested that Orrorin had independently evolved
bipedalism, separately from Lucy and other species of Australopithecus,
whom they relegated to an extinct side-branch of the hominid family.22
The controversy of Orrorin resumed again in September 2004 with the publication
of the results of computerized tomography (CT) scans of the femur, claiming a non-ape,
later hominid, distribution of cortical bone in the neck-shaft junction of Orrorin’s
femur, indicating bipedal locomotion.23
As reported by Ann Gibbons, when speaking about these latest finds:
‘Senut proposed that Orrorin’s gait was more humanlike than that of
the 2– to 4-million-year-old australopithecines. If so, australopithecines
would be bumped off the direct line to humans—a dramatic revision of our prehistory.’24
Paleoanthropologist Tim White disagreed with this ‘gait’ assessment
and attacked the latest published findings by saying that ‘the resolution
of the CT scans was so poor that it was impossible to be certain of the pattern
of bone thickness’.24 White also ‘called
Senut’s displacement of australopithecenes “une position créationniste”,
because it suggests that Orrorin’s femur was quite modern 6 million
years ago, rather than evolving by stages’.24
It seems that even evolutionists that stray outside the orthodox evolution hominid
story risk a backlash.
According to David Begun of the University of Toronto, the evidence for bipedalism
in Orrorin is ambiguous.21 Soon after
the initial publication of the fossils, Begun commented that the fossil fragments
representing Orrorin could not reveal whether it was ‘on the line
to humans, on the line to chimps, a common ancestor to both, or just an extinct
side branch’.19
Ardipithecus kadabba
A few months after Orrorin tugenensis was announced, another early hominid
candidate, from Middle Awash, Ethiopia, called Ardipithecus ramidus kadabba,
dated to between 5.2 and 5.8 Ma, was described in the journal Nature.25 The name of this specimen
was, in 2004, elevated from subspecies status to a separate species, Ardipithecus
kadabba, based on the recovery of additional fossil teeth.26 In kadabba (as in Orrorin)
the fossil finds consisted of some postcranial bones, teeth, and jaw fragments,
and so it is not known what the head of these creatures looked like.
The kadabba specimen is interpreted as a biped based on the characteristics
of a single toe bone (a proximal foot phalanx), in particular the dorsal orientation
of the proximal joint surface.27
The argument is that the toe bone’s joint surface is tilted upwards in a human-like
manner, whereas in chimpanzees it tilts downwards, and so it is supposedly evidence
that kadabba ‘toed off’ in a human-like manner when walking.28 However, as pointed out by
Begun,
‘the same joint configuration occurs in the definitely non-bipedal late Miocene
hominid Sivapithecus , and the length and curvature of this bone closely
resembles those of a chimpanzee or bonobo’.29
Amazingly, the toe bone is dated several hundred thousand years younger than the
rest of the fossils, and was found in a locality 16 km away from the rest, making
even famous hominid hunter Donald Johanson dubious about categorizing the toe bone
with the rest of the fossils.30
From the few (eleven) fossil scraps, belonging to at least five different individuals,
from five different locations,25 it seems amazing
that they can all be designated as belonging to the same subspecies, and later,
on the basis of finding some additional teeth,26
that the fossils can be assigned to a new species. According to Balter and Gibbons,
‘The Orrorin and Ardipithecus teams assert that each other’s
fossils could represent an ancestor of chimps or other apes, rather than one of
our early human ancestors or cousins.’31
Perhaps both teams are partially right, as there is little doubt that both hominids
were mere apes.
Ardipithecus ramidus
In 1994 a team led by Tim White announced the finding of Australopithecus ramidus
from Aramis, Ethiopia, which, at the time, being dated at 4.4 Ma, was the earliest
‘hominid’ find of any significance.32
About eight months later, in a corrigendum, the authors reclassified the fossils
into an entirely new genus, Ardipithecus, with the new species name being
Ardipithecus ramidus.33
The original article was accompanied by a cover photo, in the 22 September 1994
issue of Nature, of a small jaw fragment from Ardipithecus ramidus,
containing an embedded tooth, with the title ‘Earliest hominids’. Hence,
one would have expected a significant find, but instead the fossils, comprising
postcranial, dental and cranial scraps, in the authors own words, ‘shows a
host of characters usually associated with modern apes’.34
The authors argue for hominid status mainly based on a more incisiform canine morphology
and a more anterior position of the foramen magnum, as they believe ‘Acquisition
of these states at Aramis may correlate with bipedality although this remains to
be demonstrated.’35 Even
in the evolution community, not everyone is convinced that ramidus is a
hominid. According to Peter Andrews, of London’s Natural History Museum, the
thin enamel on the teeth of ramidus ‘is more of what you’d
expect from a fossil chimp’, and the features of an upper arm bone ‘suggests
knuckle-walking, chimp-style’.36
Recently, more fossil scraps from Afar, Ethiopia (jaws, teeth, toe and finger bones),
were assigned to at least nine ramidus individuals, and claims of bipedality
were made based on the dorsal orientation of the proximal joint surface of a proximal
foot phalanx, a feature also seen in kadabba.37
See the kadabba section above for a refutation of this bipedality claim.
Australopithecus anamensis
In 1995, dental, cranial and postcranial specimens from two separate localities
in Kenya, dated from about 3.8 to 4.2 Ma, were announced as belonging to a new hominid
species, Australopithecus anamensis.38
Most of the fossil scraps undoubtedly came from an ape, such as the chimp-like jaws,
but controversy has surrounded the alleged more human-like nature of the tibia and
humerus.39 The anamensis
humerus lacks a deep, oval hollow, used as a locking mechanism between the humerus
and ulna, the latter being present in chimpanzees, but not in humans, and the anamensis
tibia is wide, as in humans, because of extra spongy tissue, which acts as
shock absorbers during bipedal locomotion.40
At the time of publication, paleontologist Peter Andrews raised the suggestion that
the tibia and humerus, coming from different sections (upper level) of the Kanapoi
locality strata than the ‘primitive’ jaws and teeth (lower level), might
possibly ‘be related to humans and the other to apes’.41 There were some mandible fragments (KNM-KP
29287) found in upper-level strata, but these were from a different (higher) level
than the tibia ( KNM-KP 29285) and humerus (KNM-KP 271).42
At the time there was also the problem of several hundred thousand years in evolutionary
time between the upper and lower levels, but in a later paper, dates were obtained
that narrowed the time gap.43
In this later article additional finds of fossil scraps from anamensis
were reported, but no additional evidence was provided associating them with the
tibia and humerus. Also, the 2000 analysis of a fossil radius (KNM-ER 20419) from
the other anamensis location, Allia Bay, indicated ‘specialized wrist
morphology associated with knuckle-walking’.44
In a paper co-authored by Meave Leakey, anamensis is said to be very like
afarensis, postcranially, with the humerus, tibia and radius ‘almost
exactly matched in size and morphology to the A. afarensis collections
from Hadar’.45 Hence, if
the postcranial remains do belong to anamensis, then at most you have an
ape-like creature with a similar locomotion pattern to the ape-like afarensis.
As more in-depth analysis has been performed on afarensis, which has more
postcranial fossils attributed to it, the reader is referred to that section.
Australopithecus afarensis

The Homo erectus skeleton of KNM-WT 15000 (left) is compared to the australopithecus
afarensis skeleton of Lucy (right). There is a huge morphological gap between
the essentially modern human postcranial skeleton of erectus and that of the australopithecine
postcranial skeleton, as represented by Lucy. For example, the conical-shaped rib
cage of Lucy suggests she was thick-waisted with a pot-bellied abdomen, like chimpanzees.
Also, there is wide flaring of the ilia of Lucy’s pelvis, compared to the
narrower ilia of humans. For further discussion, see Mehlert.113
(Image from Walker and Shipman.114)
|
The discovery of the famous Lucy skeleton in Ethiopia in 197446
and the controversy over the naming of Australopithecus afarensis in 197847 are well known and will not
be retold here. The estimated cranial capacity of afarensis, whose evolutionary
time range was from about 3.0 to 3.9 Ma, is between 375 and 540 cm3,
with a mean of about 470 cm3,48
although the mean brain size is elsewhere given as 438 cm3.49 Above the neck, the skull (including jaws)
of afarensis has been described as ape-like,50
with evolutionary experts distinguishing its cranium from that of chimpanzees, mainly
due to smaller canine and larger postcanine teeth in afarensis, ‘and
the influence the smaller canines has on the face of A. afarensis, including
the reduced snout and the presence of a canine fossa’.51
Details of the first skull of afarensis (AL 444–2) were published
in 1994,52 and at the time it
was the largest australopithecine skull known,53
with an estimated cranial capacity later determined to be 540 cm3.50
Previous to this the reconstruction of the afarensis skull was a composite,
based on fossil fragments from several individuals,54
and as a result there were accusations that Kimbel and White, who made the composite
skull, ‘had force-fit the face of a slender Homo-like species onto
the brain case of a robust one’.55
Postcranially, afarensis possessed ape-like features such as a conical-shaped
rib cage, and, along with other australopithecines, were heavily built for their
stature and ‘almost certainly were not adapted to a striding gait and running,
as humans are’.56 Humans
have barrel-shaped rib cages, and by comparison to apes, are lightly built for their
stature. Analysis of the socket of the shoulder blade (scapula) of afarensis
showed that it was directed far more cranially (skywards) compared to humans,57 whose socket is perpendicular to the ground,
and that, like apes, this upward orientation of the afarensis socket would
have been ‘valuable if the arm were held overhead much of the time, as it
is when climbing and hanging in trees’.58
As for limb proportions, the estimated humerofemoral index (ratio of humerus to
femur length) in afarensis is less than that of extant apes, although still
significantly greater than that of humans.59
Humans have relatively short arms and very long legs, and according to Tattersall
and Schwartz, it is a ‘fact that the legs of afarensis were quite
short’.60 The estimated
brachial index (ratio of radius to humerus length) in afarensis is greater
than that of humans and gorillas, but less than that of other extant apes, but this
estimated index is very unreliable because of the difficulty in making a reliable
length estimate of the radius of Lucy (AL 288–1), and a generous length estimate
of the radius would bring the brachial index within the chimpanzee range.61 However, Tattersall and Schwartz state that
‘even if the relative proportions of the arms are more humanlike than apelike
(the lower arm is much longer compared with the upper arm in chimpanzees and especially
the full-time arborealists—orangutans and gibbons)—the arms of afarensis
were still quite long’.60
The bones of the fingers in afarensis, particularly the proximal phalanges,
are curved, similar to those of chimpanzees, and ‘indicate adaptation for
suspensory and climbing activities which require powerful grasping abilities’.62 The toes of afarensis
are also curved,63 as in apes,
and suggest an ability to climb trees. According to Stern and Susman, the foot and
ankle remains of afarensis ‘reveal to us an animal that engaged in
climbing as well as bipedality’.64
They go on to state that ‘There is no evidence that any extant primate has
long, curved, heavily muscled hands and feet for any purpose other than to meet
the demands of full or part-time arboreal life.’64
The analysis by Stern and Susman, in 1983, of afarensis hand fossils indicated
many features in common with chimpanzees, but they pointed out that examination
of the metacarpals yielded no evidence that afarensis was a knuckle-walker.65 Evidence for afarensis
being a knuckle-walker would surface later, however. The wrist morphology in extant
knuckle-walking African apes (chimpanzees and gorillas) has a specialized locking
mechanism that limits ‘wrist extension during the support phase of knuckle-walking’.44 In 2000 Richmond and Strait reported that the
distal radial morphology (including the scaphoid notch) in anamensis and
afarensis was similar to that of the knuckle-walking African apes, whereas
the radius attributed to robustus, and in particular africanus,
was more human-like.66 Amazingly,
the authors suggested that the knuckle-walking features in anamensis and
afarensis were non-functional retentions from an earlier ancestor.67 However, non-functional retentions (evolutionary
baggage) are not something that is expected to persist for long, according to evolutionist
Jeremy Cherfas:
‘Everything about her skeleton, from fingertips to toes, suggests that Lucy
and her sisters retain several traits that would be very suitable for climbing in
trees. Some of those same tree-climbing adaptations can still be detected, albeit
much reduced, in much later hominids such as the 2-million-year-old specimens of
Homo habilis from the Olduvai gorge. It could be argued that Lucy’s
arboreal adaptations are just a hangover from her tree-dwelling past, but animals
do not often retain traits that they do not use, and to find those same features
in specimens 2 million years later makes it most unlikely that they are remnants.’68

Lucy is the most famous Australopithecus afarensis specimen, with about
40% of the postcranial skeleton recovered.
|
Another point of interest from the Richmond and Strait study was that the wrist
morphology in africanus was more ‘modern’ than that of afarensis.
However, other studies have indicated that the foot, lower leg and limb proportions
in africanus are more ape-like than in afarensis, although the
skull of afarensis is more chimpanzee-like than africanus.69 One might well wonder what the supposed evolutionary
relationship between africanus and afarensis is in this complicated
picture, as afarensis has been proposed as the ancestor of africanus.
According to Collard and Aiello:
‘It is no longer a case of the skull pointing to one set of phylogenetic relationships,
and the postcranial skeleton—everything but the skull—to another. Rather, different
parts of the postcranium may not support the same phylogenetic hypothesis.’69
In regards to Lucy, according to archaeology professor Clive Gamble:
‘The anatomy shows a fully bipedal gait but with some differences from modern
humans. Indeed, these are so distinctive that in his exhaustive study of the limbs
Charles Oxnard concluded that the method of locomotion was neither ape-like nor
human. Nor was it a midway stage but truly unique.’70
The central argument for afarensis being a hominid is that the creature
had an erect posture and was capable of bipedal locomotion. The argument that afarensis
was bipedal is based on skeletal reconstructions of the pelvis and lower limb
bones. As up to 40% of Lucy’s postcranial skeleton was recovered, a lot of
the morphological and biomechanical analyses are based on these bones, although
not exclusively so. One problem with reconstructions of Lucy’s skeleton (dated
to about 3.2 Ma) is the subjective nature of the work, as predetermined belief about
the posture of afarensis may bias the reconstruction of the skeleton. How
else can the following statement by Maurice Abitbol, who studied Lucy’s pelvis,
make sense?
‘Prevailing views of Lucy’s posture are almost impossible to reconcile.
When one looks at the reconstruction proposed by Lovejoy (1998) and by Weaver et
al. (1985), one gets the impression that her fleshed reconstruction would
be the body of a perfectly modern human biped (Figure 1a). But when one looks at
the preliminary reconstruction recently shown at the Smithsonian, one gets the impression
of a chimpanzee awkwardly attempting to stand on its hindlimbs and about to fall
on its frontlimbs (Lewin, 1988). In the latter, the implication is a “primitive”
form of bipedality in the Hadar hominids. To resolve such differences, more anatomical
(fossil) evidence is needed. The available data at present are open to widely different
interpretations.’71
Amazingly, Lucy’s posture can be interpreted to be anything from modern human
to chimpanzee-like. Hence, creationists have every right to be extremely sceptical
of claims that Lucy and other australopithecines were at a stage of evolution between
human and apes, when the main evidence for that claim is based on such speculative
interpretation of data. From his study, Abitbol concluded that ‘Lucy’s
erect posture is unlike that seen in modern humans and is still a mystery’.72
Another problem with studies that investigate the locomotion capability of fossils
belonging to creatures such as afarensis is that discussions about muscular
functions ‘assume a priori that the muscles were in a close to human
pattern’.73 In a biomechanical
analysis of Lucy’s pelvis and lower limb bones, in particular the reconstruction
of the gluteal musculature, Christine Berge reported that ‘an ape-like
gluteal organization would offer better ability than a human-like one’.73
From the study, Berge concluded
‘that the bipedalism of Australopithecus must have differed from
that of Homo. Not only did Australopithecus have less ability
to maintain hip and knee extension during the walk, but also probably moved the
pelvis and lower limb differently. It seems that the australopithecine walk differed
significantly from that of humans, involving a sort of waddling gait, with large
rotatory movements of the pelvis and shoulders around the vertebral column (Berge,
1991a, b). Such a walk, likely required a greater energetic cost than does human
bipedalism.’74
From their detailed analysis of the postcranial skeleton, Stern and Susman ‘discovered
a substantial body of evidence indicating that arboreal activities were so important
to A. afarensis that morphologic adaptations permitting adept movement
in trees were maintained’.75
Concerning afarensis, they also concluded ‘that the nature of terrestrial
bipedality, when it was practiced, was different from modern humans’.75
Also, CT scans by a group led by anatomist Fred Spoor of the University of Liverpool
of the bony labyrinth (fluid-filled semicircular canals) of the inner ear, which
is part of the vestibular apparatus responsible for balance,76
showed that the semicircular canal dimensions in crania attributed to other australopithecines
(africanus and robustus) were similar to that of extant great
apes.77 Although scans of afarensis
were not performed, it is likely that the labyrinths of afarensis
would resemble the same chimp/ape pattern as these other australopithecines.78 Hence, the evidence indicates that it is
very unlikely that afarensis was a habitual bipedal walker. It is also
not clear whether the limited bipedalism exhibited by some australopithecines, such
as afarensis, was postural and related to feeding, as opposed to locomotion.79 It should be remembered that
extant bonobos (pygmy chimpanzees) have been observed to walk bipedally about 10%
of the time, so limited non-human bipedalism occurs even in extant apes.79
So what exactly do we have in afarensis? The answer is a creature with
the brain the size of an ape, a skull that was ape-like with a body similar in shape
and size to an ape, and a creature that was specialized for climbing in trees and
knuckle walking, similar to apes. In other words, you have an ape. However, if afarensis
also had some limited ability for non-human bipedal locomotion, which was perhaps
more efficient than that of extant bonobos, is this evidence that the creature was
an apeman, or does it just reflect the diversity of the ape/australopithecine body
structure that God created?
That the answer lies with the latter is indicated in the analysis of the skeletal
remains of an ape from Italy called Oreopithecus bambolii, dated from 7
to 9 Ma, whom nobody appears to want to label a hominid, as the fossil remains are
in the wrong location and outside the hominid time ‘window’, yet bambolii
had an anatomy supporting limited bipedal locomotion.80
According to the authors of the study, parts of the pelvis of bambolii
resembled that of afarensis, and its femur showed ‘a pronounced diaphyseal
angle combined with condyles of subequal size, similar to Australopithecus
and Homo and functionally correlated with bipedal activities’. 81 According to Henry Gee, ‘this
creature is thought to have become bipedal independently and was only distantly
related to hominids’.82
Apes evolving a form of bipedal locomotion once is difficult enough to believe or
imagine; that it must have independently happened multiple times, in order to ‘rescue’
evolutionary theory, reveals evolution to be a collection of just-so stories that
can be accommodated to almost any scenario, no matter how absurd or unlikely. That
a limited non-human form of bipedal locomotion existed in apes unrelated to, and
earlier than, any hypothetical hominid, indicates that possession of bipedal traits
are not indicative of hominid status.
Kenyanthropus platyops
In March 2001 yet another ‘hominid’ appeared on the cover of Nature,
with the headlines reading ‘The human family expands’. Inside was an
article describing a new creature, from Lake Turkana, Kenya, called Kenyanthropus
platyops.83 Being dated
at 3.5 Ma has brought platyops into direct competition with afarensis
as the trunk hominid that supposedly gave rise to the genus Homo.84 The main excitement was over a badly distorted
cranium. Its cranial capacity was estimated to be of ape proportions, but largely
because of its flat face, unusual for a supposed hominid at that stage of evolution,
the specimen was assigned to a new genus Kenyanthropus. However, there
are strong indications that the flat face was due to severe expanding matrix distortion,
an artifact of the fossilization process,85
and so platyops is undoubtedly just another ape.
Australopithecus bahrelghazali
In 1995 the mandible from this creature, found in Chad, was described as being most
similar to the contemporary species afarensis.86
The specimen from Chad, date estimated at between 3 and 3.5 Ma, was, in 1996, assigned
to the new species Australopithecus bahrelghazali. It was judged different
from afarensis on certain morphological features of the mandible, premolar
roots and premolar enamel thickness. However, such a meager find hardly warrants
the creation of a new species. There is no reason to view bahrelghazali
as anything but an ape, probably of the same kind as afarensis.
Australopithecus africanus

Photo by David Green
The Australopithecus africanus cranium STS 5 (nicknamed ‘Mrs Ples’)
is one of the most complete australopithecine crania ever found. It was discovered
in 1947 by Robert Broom and John Robinson at Sterkfontein, South Africa. It has
a cranial capacity of about 485 cm3.
|
The mean brain size of seven specimens of Australopithecus africanus is
451 cm3, with a range from 425 cm3 to 515 cm3.87 Evolutionists believe that
africanus lived in South Africa from about 2.5 to 3 Ma, although it may
have persisted until 2 Ma.88 Cranially,
the main difference in comparison to afarensis is in the face, with the
face of africanus less prognathic and broader.89
Also, in comparison to afarensis, the africanus lower jaw has
a more robust body, the postcanine teeth are enlarged in size, and the anterior
teeth are reduced.89
Australopithecus afarensis is considered by many as being ancestral to
africanus, but comparisons of the limb proportions of the two species indicate
less ape-like forelimb to hindlimb joint sizes and limb lengths in the supposedly
older and craniodentally more ‘primitive’ afarensis.90 According to the authors of these studies,
‘This implies that limb proportions changed back and forth in the hominid
lineage or our present view of hominid relationships is too simplified.’91 Rather, the confusion of hominid
relationships exists because there never was an evolutionary relationship between
hominids to begin with. According to Klein and Edgar, africanus possessed
a very ape-like upper body, with long, powerful arms suitable for tree climbing,
but the lower body was ‘shaped for habitual bipedal locomotion on the ground’.92 However, as mentioned above,
CT scans of the bony labyrinth of the inner ear showed that the semicircular canal
dimensions in crania attributed to africanus were similar to those of extant
great apes.77 Hence, if africanus did
walk bipedally in some manner, it is unlikely to have been habitually.
An earlier analysis of the postcranial anatomy of afarensis and africanus
led McHenry to the conclusion
‘that (1) the postcranial skeleton of early Australopithecus was
uniquely different from all living apes and people, (2) the postcranial skeleton
of early A. africanus is very similar to A. afarensis despite
real differences in their skulls, teeth, distribution, and age, and (3) there is
a distinct grade of locomotor adaptation in Australopithecus which is unlike
any living form.’93
Hence, if the two species shared ‘a unique configuration which implies that
locomotor and postural adaptations were very similar and unlike any living form’,94 then the locomotor behaviour
of africanus was similar to afarensis. As indicated earlier, afarensis
was a knuckle-walking ape built for an arboreal lifestyle, although it may
have had a limited ability to walk bipedally in a non-human-like manner, as did
apes unrelated to any supposed hominid evolution, such as Oreopithecus bambolii.
As with all other australopithecine species, africanus possessed brains
the size of apes, had ape-like skulls, and was similar in body shape and size to
apes. Hence, the most plausible explanation is that africanus was a
species of extinct ape, and even among evolutionists there have been suggestions
that africanus and afarensis ‘should be considered subspecies
of a single species’.95
Australopithecus garhi
In 1999 the remains of Australopithecus garhi, a supposed 2.5-million-year-old
hominid from the Hata beds of Ethiopia’s Middle Awash, was said to be distinguished
from afarensis ‘by its absolutely larger postcanine dentition and
an upper third premolar morphology with reduced mesiobuccal enamel line projection
and less occlusal asymmetry’.96
However, when viewing the very incomplete cranial remains of the creature, with
a cranial capacity estimated at 450 cm3, it is difficult not to wonder
whether differences in a few dental features have been overemphasized, as this specimen
surely could have been assigned to the species afarensis. Perhaps the later
evolutionary date was a factor. Regardless of its affinity to afarensis,
the ape-like nature of its skull establishes garhi as nothing more than
an ape.
More interesting were the postcranial fossils (BOU-VP-12/1) recovered at the locality,
which the researchers did not assign to garhi, featuring a human-like humerofemoral
index.97 However, the Bouri skeleton
(BOU-VP-12/1) also had an estimated brachial index larger than all extant apes,
except orangutans (Pongo pygmaeus).98
This combination of a long, ape-like forearm and human-like humerofemoral limb proportions
makes for a confusing ancestor–descendant evolutionary relationship, as it ‘would
suggest a reversal in brachial proportions in the Bouri skeleton (more ape-like)
from the A. afarensis condition, followed by a second reversal in H. ergaster
(more human-like)’.99
The robust australopithecines
The robust australopithecines are conventionally made up of the species Australopithecus
robustus (1.5 to 1.8 Ma) from South Africa, and the two East African species
Australopithecus boisei (1.3 to 2.3 Ma) and Australopithecus aethiopicus
(2.3 to 2.7 Ma), with the combined evolutionary age range for the three species
from 1.3 to 2.7 Ma.100 Some
paleoanthropologists assign them to a separate genus, Paranthropus. None
of the robust australopithecine species are regarded by evolutionists as ancestors
of true humans, but rather as side branches that met a dead end, 101 ‘because their teeth and skulls were
so specialized and because they coexisted with more plausible ancestors after 2.5
million years ago’.102
It is because of ‘their huge chewing teeth and rugged skulls’ that they
have been called robust australopithecines, although they had small brains and bodies.103 From the skulls of these
creatures, it is obvious that there was nothing human-like about them.
Australopithecus aethiopicus
The famous ‘Black Skull’ (KNM-WT 17000), with a cranial capacity of
about 410 cm3, is one of the three or four fossil specimens making up
the species aethiopicus,104
which is considered by some evolutionists to be a plausible link between afarensis
and the other two robust species, boisei and robustus.105 However, the Black Skull ‘is an extreme
A. robustus/boisei type with a massive sagittal crest, and the two teeth
found are four or five times as large as human molars’,106
and so even though aethiopicus is supposedly an older species then robustus/boisei,
its hyper-robustness disqualifies it as a transitional form. The morphology of the
Black Skull has many similarities to that of a small male gorilla,107 and so it is possible that gorillas and
aethiopicus belong to the same biblical kind. There are currently no postcranial
fossils assigned to aethiopicus, the other two (possibly three) specimens
attributed to aethiopicus being mandibles.108
Australopithecus boisei
The mean brain size of Australopithecus boisei, based on six specimens,
is about 452 cm3, with a range from 390 cm3 to 500 cm3,109 although other estimates
of boisei give a mean of 521 cm3.49
Cranially, boisei is described as ‘the only hominin that combines
a massive, wide, flat, face with a modest-sized neurocranium’.110 There are no postcranial remains that can
definitely be attributed to boisei, but from Koobi Fora a partial skeleton
and individual postcranial bones have been inconclusively linked with the species.110 According to Wood and Richmond, ‘The
partial skeleton is characterised by limb proportions that resemble A. afarensis
(Grausz et al. 1988), and are less apelike than those of A. africanus’.110 Hence, if the partial skeleton (KNM-ER 1500)
did belong to boisei, then it may have had similar locomotor behaviour
to that of afarensis, and ‘like A. afarensis, the relatively
long forelimb suggests that the locomotor behaviour of P. boisei included
an arboreal component’.111
Australopithecus robustus

Photo by Warwick Armstrong
As can be seen from this replica skull of Australopithecus boisei, there
was nothing human about the heads of the robust australopithecines. In a similar
replica at the San Diego Museum of Man, a lower jaw has been included, which is
based on a fossil found 50 miles away.
|
The brain size of Australopithecus robustus, which currently can only be
estimated for one specimen (SK 1585), was 530 cm3, but a more recent
estimate has lowered that to 476 cm3.109
Cranially, robustus was similar to that of boisei, but its face
was not as large and wide.110 Hence, if robustus
had a ‘robust’ skull, then the skull of boisei was hyper-robust.
In comparison to africanus, the chewing teeth and face of robustus
were larger, although its canine and incisor teeth were smaller,112 and according to Klein and Edgar, both
species
‘possessed very ape-like upper bodies with long, powerful arms that would
have made them agile tree climbers. They differed from apes primarily in their lower
bodies, which were shaped for habitual bipedal locomotion on the ground, and in
their teeth’.92
According to Wood and Richmond, the morphology of the pelvis and hip believed to
be associated with robustus resembles that of afarensis and africanus
(the ‘gracile’ australopithecines), and suggests that ‘the
gait of P. robustus probably resembled that of the ‘gracile’
australopiths’.112 As indicated earlier,
the gait of afarensis was not human-like. Also, as mentioned previously,
CT scans of the bony labyrinth of the inner ear showed that the semicircular canal
dimensions in crania attributed to robustus were similar to that of extant
great apes.77
Conclusion
The morphological features of the hominid fossils assigned to taxa outside the genus
Homo indicate they were mere extinct apes. A species is not equivalent
to a biblical kind, and several of the non-Homo hominids may just
represent variation within one or more ape kinds, just as the fossil species
included in Homo, excluding the invalid taxon Homo habilis, are
believed to represent variation within the one human kind. It may well
be that extant apes are surviving remnants of these hominid ape kinds,
which have lost much of their genetic diversity. In general, where fossil material
is available, these hominid creatures had brains the size of apes, skulls that were
ape-like, and bodies similar in shape and stature to apes. The morphology of creatures
such as Australopithecus afarensis indicates that they were specialized
for climbing in trees, as well as knuckle walking, as are apes. Much has been made
of skeletal features, indicating some of the australopithecines, including afarensis,
may also have had limited ability for non-human bipedal locomotion. However, similar
limited bipedal ability also existed in apes not considered hominids, such as Oreopithecus
bambolii, dated to earlier than the supposed human and chimpanzee split.
Hence, how can a trait be used as an argument for the uniqueness of all hominids,
and their evolutionary relationship, when the trait is not unique to these supposed
hominids? It is like saying whales must have evolved from creatures like hippopotamuses
because they can both swim (or do they believe that, too?). I suppose this is why
evolutionists like the word parallel so much, as in parallel evolution. Call it
what you like, but is it science?
References
- Harrub, B. and Thompson, B., The Truth About Human Origins,
Apologetics Press Inc., Montgomery, AL, pp. 8–10, 2003. Return
to Text
- Gribbin, J. and Cherfas, J., The First Chimpanzee: In Search
of Human Origins, Penguin Books, London, pp. 116–117, 2001.
Return to Text
- Sarfati, J., Refuting Compromise, Master Books, Green
Forest, AR, pp. 376–388, 2004; Lubenow, M.L., Bones of Contention: A
Creationist Assessment of Human Fossils, Revised and Updated, Baker Books,
Grand Rapids, MI, pp. 273–292, 2004. Return to Text
- Harrub and Thompson, ref. 1, pp. 99–134; Lubenow, ref.
3, pp. 222–235. Return to Text
- Lubenow, ref. 3, p. 187. Return to Text
- Brunet, M. et al., A new hominid from the upper Miocene
of Chad, Central Africa, Nature 418:145–151, 2002.
Return to Text
- Wood, B., Hominid revelations from Chad, Nature
418:134, 2002. Return to Text
- BBC News, Skull find sparks controversy, <news.bbc.co.uk/hi/english/sci/tech/newsid_2125000/2125244.stm>,
13 July 2002. Return to Text
- Montgomery, B., Man’s ancestors 7m years old, The
Australian, 11 July 2002, p. 1. Return to Text
- Wolpoff, M.H., Senut, B., Pickford, M. and Hawks, J., Sahelanthropus
or ‘Sahelpithecus’? Nature 419:581–582,
2002. Return to Text
- Ancient skull more ape than human? <www.cnn.com/2002/TECH/science/10/09/ancient.skull/index.html>,
10 October 2002. Return to Text
- Wolpoff et al., ref. 10, p. 582.
Return to Text
- Brunet et al., ref. 6, p. 151.
Return to Text
- Murdock, M., Sahelanthropus tchadensis—the ambiguous
ape, TJ 18(3):114, 2004. Return to Text
- Beauvilain, A. and Guellec, Y.L., Further details concerning
fossils attributed to Sahelanthropus tchadensis (Toumai), South African
J. Science 100:142–144, 2004. Return
to Text
- Beauvilain and Guellec, ref. 15, p. 142.
Return to Text
- Dalton, R., Brickbats for fossil hunter who claims skull has
false tooth, Nature 430:956, 2004. Return
to Text
- Aiello, L.C. and Collard, M., Our newest oldest ancestor?
Nature 410:526–527, 2001. Return to
Text
- Balter, M., Scientists spar over claims of earliest ancestor,
Science 291:1460, 2001. Return to Text
- Wong, K., An ancestor to call our own, Scientific American
Special 13(2):9, 2003. Return to Text
- Begun, D.R., The earliest hominins—is less more?
Science 303:1479, 2004. Return to Text
- Shipman, P., Hunting the first hominid, American Scientist
90(1), 2002; <www.americanscientist.org/template/AssetDetail/assetid/18641>,
13 August 2004. Return to Text
- Galik, K., Senut, B., Pickford, M., Gommery, D., Treil, J.,
Kuperavage, A.J. and Eckhardt, R.B., External and internal morphology of the BAR
1002'00 Orrorin tugenensis femur, Science 305:1450–1453,
2004. Return to Text
- Gibbons, A., Oldest human femur wades into controversy,
Science 305:1885, 2004. Return to Text
- Halle-Selassie, Y., Late Miocene hominids from the Middle Awash,
Ethiopia, Nature 412:178–181, 2001.
Return to Text
- Halle-Selassie, Y., Suwa, G. and White, T.D., Late Miocene
teeth from Middle Awash, Ethiopia, and early hominid dental evolution, Science
303:1503–1505, 2004. Return to Text
- Halle-Selassie, ref. 25, p. 180. Return
to Text
- Wong, ref. 20, pp. 8–9. Return to
Text
- Begun, ref. 21, pp. 1478–1480. Return
to Text
- Lemonick, M.D. and Dorfman, A., One giant step for mankind,
Time (South Pacific) pp. 58–59, 23 July 2001. Return
to Text
- Balter, M. and Gibbons, A., Another emissary from the dawn
of humanity, Science 293:189, 2001. Return
to Text
- White, T.D., Suwa, G. and Asfaw, B., Australopithecus ramidus,
a new species of early hominid from Aramis, Ethiopia, Nature 371:306–312,
1994. Return to Text
- White et al., ref. 32, p. 88.
Return to Text
- White et al., ref. 32, p. 311.
Return to Text
- White et al., ref. 32, p. 312.
Return to Text
- Fischman, J., Putting our oldest ancestors in their proper
place, Science 265:2011, 1994. Return
to Text
- Semaw, S. et al., Early Pliocene hominids from Gona,
Ethiopia, Nature 433:304, 2005. Return
to Text
- Leakey, M.G., Feibel, C.S., McDougall, I. and Walker, A., New
four-million-year-old hominid species from Kanapoi and Allia Bay, Kenya, Nature
376:565–571, 1995. Return to Text
- Leakey, M. and Walker, A., Early hominid fossils from Africa,
Scientific American 276(6):63–64, 1997.
Return to Text
- Leakey and Walker, ref. 39, p. 63. Return
to Text
- Andrews, P., Ecological apes and ancestors, Nature
376:556, 1995. Return to Text
- Leakey, M.G., Feibel, C.S., McDougall, I., Ward, C. and Walker,
A., New specimens and confirmation of an early age for Australopithecus anamensis,
Nature 393:62, 1998; The distal end of the humerus was
discovered in 1965 and is described in: Patterson, B. and Howells, W.W., Hominid
humeral fragment from early Pleistocene of Northwest Kenya, Science
156:64–66, 1967. Return to Text
- Leakey et al., ref. 42, p. 62.
Return to Text
- Richmond, B.G. and Strait, D.S., Evidence that humans evolved
from a knuckle-walking ancestor, Nature 404:382, 2000.
Return to Text
- Ward, C., Leakey, M. and Walker, A., The new hominid species
Australopithecus anamensis, Evolutionary Anthropology 9:201,
1999. Return to Text
- Reader, J., Missing Links: The Hunt for Earliest Man,
Penguin Books, London, pp. 207–210, 1988. Return to Text
- Lewin, R., Bones of Contention, Penguin Books, London,
pp. 268–300, 1987. Return to Text
- Wood, B. and Richmond, B.G., Human evolution : taxonomy and
paleobiology, J. Anatomy 196:29, 2000.
Return to Text
- McHenry, H.M. and Coffing, K., Australopithecus to
Homo: transformations in body and mind, Annual Review of Anthropology
29:127, 2000. Return to Text
- Tattersall, I. and Schwartz, J.H., Extinct Humans,
Westview Press, New York, p. 88, 2001. Return to Text
- Wood and Richmond, ref. 48, p. 28. Return
to Text
- Kimbel, W.H., Johanson, D.C. and Rak, Y., The first skull and
other new discoveries of Australopithecus afarensis at Hadar, Ethiopia,
Nature 368:449–451, 1994. Return
to Text
- Aiello, L.C., Variable but singular, Nature
368:399, 1994. Return to Text
- Kimbel, W.H., White, T.D. and Johanson, D.C., Cranial morphology
of Australopithecus afarensis : a comparative study based on a composite
reconstruction of the adult skull, American J. Physical Anthropology
64:337–388, 1984; Kimbel, W.H. and White, T.D., A revised reconstruction
of the adult skull of Australopithecus afarensis, J. Human Evolution
17:545–550, 1988. Return to Text
- Shreeve, J., ‘Lucy,’ crucial early human ancestor,
finally gets a head, Science 264:34, 1994.
Return to Text
- Leakey, R. and Lewin, R., Origins Reconsidered: In Search
of What Makes us Human, Abacus, London, pp. 193–196, 1992.
Return to Text
- Stern, J.T. and Susman, R.L., The locomotor anatomy of
Australopithecus afarensis, American J. Physical Anthropology
60:284–285, 1983. Return to Text
- Cherfas, J., Trees have made man upright, New Scientist
97:172, 1983. Return to Text
- Richmond, B.G., Aiello, L.C. and Wood, B.A., Early hominin
limb proportions, J. Human Evolution 43:534–535,
2002. Return to Text
- Tattersall and Schwartz, ref. 50, p. 90.
Return to Text
- Richmond et al., ref. 59, pp. 542–543. Return to Text
- Stern and Susman, ref. 57, p. 284. Return
to Text
- Stern and Susman, ref. 57, p. 307. Return
to Text
- Stern and Susman, ref. 57, p. 308. Return
to Text
- Stern and Susman, ref. 57, pp. 283–284.
Return to Text
- Richmond and Strait, ref. 44, p. 383.
Return to Text
- Richmond and Strait, ref. 44, pp. 383–384.
Return to Text
- Cherfas, ref. 58, p. 176. Return to Text
- Collard, M. and Aiello, L.C., From forelimbs to two legs,
Nature 404:339, 2000. Return to Text
- Gamble, C., Timewalkers: The Prehistory of Global Colonization,
Penguin Books, London, p. 58, 1993. Return to Text
- Abitbol, M.M., Lateral view of Australopithecus afarensis:
primitive aspects of bipedal positional behavior in the earliest hominids, J. Human
Evolution 28:228, 1995. Return to Text
- Abitbol, ref. 71, p. 211. Return to Text
- Berge, C., How did australopithecines walk? A biomechanical
study of the hip and thigh of Australopithecus afarensis, J. Human Evolution
26:270, 1994. Return to Text
- Berge, ref. 73, p. 271. Return to Text
- Stern and Susman, ref. 57, p. 313. Return
to Text
- Shipman, P., Those ears were made for walking, New Scientist
143:27, 1994. Return to Text
- Spoor, F., Wood, B. and Zonneveld, F., Implications of early
hominid labyrinthine morphology for evolution of human bipedal locomotion, Nature
369:645, 1994. Return to Text
- Gibbons, A., Anthropologists take the measure of humanity,
Science 264:350, 1994. Return to Text
- Lubenow, M.L., Paleoanthropology in review, TJ
10(1):13, 1996. Return to Text
- Kohler, M. and Moya-Sola, S., Ape-like or hominid-like? The
positional behavior of Oreopithecus bambolii reconsidered, Proc. Nat. Acad.
Sci. USA 94:11747–11750, 1997. Return
to Text
- Kohler and Moya-Sola, ref. 80, p. 11747.
Return to Text
- Gee, H., Return to the planet of the apes, Nature
412:131, 2001. Return to Text
- Leakey, M.G., Spoor, F., Brown, F.H., Gathogo, P.N., Klarie,
C., Leakey, L.N. and McDougall, I., New hominin genus from eastern Africa shows
diverse middle Pliocene lineages, Nature 410:433–440,
2001. Return to Text
- Wong, K., Finding Homo sapiens’ lost relatives,
Scientific American 285(4):26–27, 2001.
Return to Text
- White, T., Early hominids—diversity or distortion?
Science 299:1994–1997, 2003. Return
to Text
- Brunet, M., Beauvilain, A., Coppens, Y., Heintz, E., Moutaye,
A.H.E. and Pilbeam, D., The first australopithecine 2,500 kilometres west of the
Rift Valley (Chad), Nature 378:273–275, 1995. Return to Text
- Falk, D., Redmond Jr, J.C., Guyer, J., Conroy, G.C., Recheis,
W., Weber, G.W. and Seidler, H., Early hominid brain evolution: a new look at old
endocasts, J. Human Evolution 38:709, 2000.
Return to Text
- Klein, R.K. and Edgar, B., The Dawn of Human Culture,
John Wiley & Sons Inc., New York, p. 35, 2002. Return to Text
- Wood and Richmond, ref. 48, p. 32. Return
to Text
- McHenry, H.M. and Berger, L.R., Body proportions in Australopithecus
afarensis and A. africanus and the origin of the genus Homo,
J. Human Evolution 35:1–22, 1998; McHenry, H.M.
and Berger, L.R., Limb lengths in Australopithecus and the origin of the
genus Homo, South African J. Science, 94 :447–450,
1998. Return to Text
- McHenry and Berger, ref. 90, p. 447. Return
to Text
- Klein and Edgar, ref. 88, p. 37. Return
to Text
- McHenry, H.M., The first bipeds: a comparison of the A.
afarensis and A. africanus postcranium and implications for the evolution
of bipedalism, J. Human Evolution 15:179, 1986.
Return to Text
- McHenry, ref. 93, p. 186. Return to Text
- McHenry, ref. 93, p. 187. Return to Text
- Asfaw, B., White, T., Lovejoy, O., Latimer, B., Simpson, S.
and Suwa, G., Australopithecus garhi : a new species of early hominid from
Ethiopia, Science 284:631, 1999. Return
to Text
- Asfaw et al., ref. 96, pp. 632–633.
Return to Text
- Richmond et al., ref. 59, p. 542.
Return to Text
- Richmond et al., ref. 59, p. 543.
Return to Text
- McCollum, M.A., The robust australopithecine face: a morphogenetic
perspective, Science 284:301, 1999.
Return to Text
- Johanson, D. and Edgar, B., From Lucy to Language,
Simon & Schuster Editions, New York, p.152, 1996. Return to
Text
- Klein and Edgar, ref. 88, p. 41. Return
to Text
- Klein and Edgar, ref. 88, p. 39. Return
to Text
- Johanson and Edgar, ref. 101, pp. 23, 152.
Return to Text
- Klein and Edgar, ref. 88, pp. 57–58.
Return to Text
- Mehlert, A.W., A review of the present status of some alleged
early hominds, TJ 6(1):32, 1992. Return
to Text
- Mehlert, ref. 106, p. 34. Return to
Text
- Johanson and Edgar, ref. 101, p. 23.
Return to Text
- Falk et al., ref. 87, pp. 708–709.
Return to Text
- Wood and Richmond, ref. 48, p. 37. Return
to Text
- Wood and Richmond, ref. 48, p. 38. Return
to Text
- Wood and Richmond, ref. 48, p. 35. Return
to Text
- Mehlert, A.W., Australopithecines—the extinct southern
apes of Africa: a fresh light on their status? TJ 14(3):91–99,
2000.Return to Text
- Walker, A. and Shipman, P., The Wisdom of Bones: In Search
of Human Origins, Weidenfeld and Nicolson, London, pp. 192, 1996.
Return to Text
|